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Commentary Open Access

Investigating DIX-mediated interactions in Wnt signaling with AlphaFold2 and AlphaFold3

  • 1Division of Natural and Mathematical Sciences, LeMoyne-Owen College, Memphis, TN 38126, USA
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Corresponding Author

Ho-Jin Lee, ho-jin_lee@loc.edu

Received Date: March 31, 2026

Accepted Date: July 27, 2026

Abstract

The interactions between the DIX domains of three proteins (Axin1/2, Dishevelled1/2/3, and Coiled-coil-DIX1) are essential to the mechanism of action downstream of the Wnt/β-catenin signaling pathway. Structural and biophysical studies have shown that DIX domains polymerize via head-to-tail interface interactions. In our recently published study entitled 'Exploring DIX-DIX Homo- and Hetero-Oligomers in Wnt Signaling with AlphaFold2", we examined the monomer structures of DIX domains, and DIX-mediated homodimers and heterodimers with AlphaFold2 (AF2) ColabFold. First, we evaluated the AF2-based prediction by comparing the reported monomer and complex structures of the DIX domains. The results showed that AF2 is an excellent tool for predicting the 3D structures of monomers of DIX domains and of homodimers and heterodimers of DIX-mediated proteins. Second, we evaluated the calculated binding affinities (KD) of DIX domains using the PRODIGY method. The results showed that the calculated KD values are compatible with experimentally obtained values. In this commentary, we present new computational results using AlphaFold3 (AF3), the latest version of AlphaFold, and VD-MM/GBSA calculations with HawkDock2. Overall, AF2/3 and other bioinformatics tools provide valuable insights into the DIX-mediated signaling pathway at the molecular level.

Keywords

AlphaFold2/3, HawkDock2, VD-MM/GBSA, DIX domain, Protein-Protein Interactions, Wnt signaling

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